Gramma the tortoise, the oldest living resident at the San Diego Zoo, died at the age of 141, according to the zoo.

Her life spanned a tumultuous period of U.S. and world history, from Chester Arthur to Donald Trump. To San Diegans, she was ...

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...

San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, ...

Gramma, a 141-year old Galapagos tortoise who has lived through two world wars, has passed away in a US Zoo, with generations of zoo-goers sharing their cherished memories of the beloved animal.

Gramma, one of San Diego Zoo’s most beloved residents, has died at around 141 years old, zoo officials said. She was born in ...

Zoo officials said she came from the Bronx Zoo in either 1928 or 1931 as part of their first group of Galápagos tortoises.

San Diego Zoo staff and animal lovers are reeling this week following the death of beloved Gramma the Galápagos tortoise, by ...

Tortoises can live for more than 100 years in the wild and close to double that in captivity. This tortoise was born in her ...